Correction to: Nature Genetics https://doi.org/10.1038/s41588-018-0055-6, published online 19 February 2018

In the version of this article originally published, the positions of Wenjie Shu and Qiang Wang in the author list were reversed and incorrect images were displayed in the HTML for Supplementary Figs. 1–12. The errors have been corrected in the HTML and PDF versions of the article.