Correction to: Nature Genetics https://doi.org/10.1038/s41588-019-0535-3, published online 29 November 2019.

In the version of this article initially published, an affiliation indicating that authors Anita Göndör and Rolf Ohlsson jointly supervised the work was missing. The Author contributions have additionally been updated to state that these authors contributed equally as joint senior authors. The errors have been corrected in the HTML and PDF versions of the article.